Caterpillars impact the entire food chain

The grapeleaf skeletonizer (Harrisina americana) on a Vitis vulpina (with common names frost grape, winter grape, fox grape, and wild grape.)

If you see an abundance of caterpillars on a grape leaves what is your first thought? In recent times the influence of commercials would have us reaching for a pesticide. Today through a systems theory lens I see an important character in a living system.